Key:  Family Pholadidae--Piddock clams

Phylum Mollusca
 Class Bivalvia
  Subclass Heterodonta
   Order Myoida
    Suborder Pholadina
 

Taken primarily from  Kozloff, 1987, 1996 p.284  (Copyright 1987, 1996, University of Washington Press.  Used in this web page by permission of University of Washington Press)
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Not in key:  Parapholas californica
 
1a  Rasping anterior portion of valves abruptly separated by a groove from the non-rasping portion (photo) 2
1b Rasping anterior portion of valves not abruptly separated from the non-rasping portion (burrowing in firm mud or clay; length to about 5 cm; not likely to be found north of southern Washington) Barnea (Anchomasa) subtruncata
2a Without a myophore (apophysis) in either valve; posterior portion of shell, in mature specimens, tapering to a point, thus resembling a bird's beak; anterior portion of shell gaping as if abruptly cut off; siphonoplax (consisting of a pair of flaps that enclose the siphons) calcified Netastoma rostrata
2b With a myophore (photo) in both valves; posterior portion of shell not tapering sufficiently to resemble a bird's beak (it may taper, however, especially in Pholadidea conradi); anterior portion of shell gaping (photo) (except in full-grown specimens, in which the gape is closed by a callum), but not appearing to have been abruptly cut off; siphonoplax not calcified 3
3a Rasping anterior portion of each valve occupying about half the area of the valve; valves without a shieldlike plate (protoplax) dorsal to the rasping portion; anteroventral gape not becoming closed, when the animal is full grown, by a callum; length up to 15 cm; burrowing in firm mud or clay Zirfaea pilsbryi
3b Rasping portion of each valve occupying less than half the area of the valve (photo); valves with a thick, shieldlike plate (protoplax) dorsal to the rasping portion (photo); anteroventral gape (photo) becoming closed, in full-grown specimens, by a callum; length not exceeding 8 cm; generally boring in shale, sometimes in hard clay 4
4a Posterior (siphonal) ends of valves of mature specimens with a siphonoplax (this consists of a pair of flaps that enclose the siphons); siphons smooth 5
4b Posterior ends of valves of mature specimens without a siphonoplax; siphons smooth or slightly warty 6
5a Siphonoplax consisting of flexible flaps, not lined by calcareous granules; length to about 8 cm; generally boring in shale Penitella penita
5b Siphonoplax not flexible, lined with calcareous granules; length to about 2.5 cm; generally boring in shells of abalones (Haliotis), Mytilus californianus, and other molluscs. Penitella conradi
6a Siphons smooth; umbonal reflections (calcified plates between the protoplax and the rasping portion of each valve) tightly applied to the shell (Note:  What was formerly called a protoplax on this species is now recognized as a dorsal extension of the callum--this species does not actually have a protoplax) Penitella turnerae
6b Siphons warty; umbonal reflections free of the shell for about half their length (P. gabbi in original key) Penitella richardsoni



Taxonomic Levels Represented in This Key:

  Parapholas californica
  Penitella penita
  Penitella turnerae
  Zirfaea pilsbryi



Page created by Dave Cowles, 2006
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