Other Valuable Web Resources for Studying Marine Invertebrates
in the Puget Sound and Straits of Juan de Fuca:
Marine Stations on the Pacific Coast of North America
Laboratory: Anacortes, WA, on Fidalgo
by Walla Walla College. Undergraduate and M.S. level marine
and research. Parent site for this survey of invertebrates.
In Bodega Bay, CA north of San Francisco. Operated by the
of California, Davis
In Friday Harbor, WA, on San Juan Island. Operated by the
of Washington. Marine research by graduate students and
Tiburon Center for Environmental
Studies: San Francisco State University's
center for marine
and estuarine research. Located on San Francisco
Point Marine Center:
In Anacortes, WA on Fidalgo Island. Operated by Western
University. Undergraduate marine research and classes.
Near Santa Cruz in Monterey Bay. Operated by the Institute of
Sciences at UC Santa Cruz.
Field Station: On Blakely Island, WA, one
of the San Juan
Islands. Operated by Seattle Pacific University and Seattle
The Mission of the Blakely Island Field Station is to support
in education and research in field-based environmental and physical
while supporting the preservation and wise use of Blakely Island
Landing Marine Laboratories:
In Monterey Bay, CA. Operated by California State University
Marine Science Center:
On the Yaquina Bay estuary In Newport, Oregon. Operated by
State University. A research and teaching facility.
Bay Aquarium Research Institute:
In Monterey Bay, CA. An independent institution affiliated
Monterey Bay Aquarium. Established by David Packard of the
Foundation. Studies on biology and engineering in the deep
of Marine Biology: Charleston, Oregon
near Coos Bay.
Operated by the University of Oregon. A research and teaching
In Pacific Grove, CA. Operated by Stanford University
Is on the campus of the University of California, Santa Barbara.
Preserve: A marine preserve near
Victoria, BC, Canada, on
the grounds of the old Race Rocks light station. Sponsored by
Lester B. Pearson College, which offers the International Baccalaureate
Wrigley Institute for Environmental Studies:
At Two Harbors
on Catalina Island, off Los Angeles, CA. Operated by the
of Southern California. Studies marine and environmental
Marine Sciences Center:
Bamfield, BC, Canada, on Barkley Sound on the protected side of
Island. Operated by the Western Canadian Universities Marine
Society. Includes a Biodiversity
database recording species found near the station.
Kerckhoff Laboratory: In Newport Beach,
by California Institute of Technology. Focus mainly on
and developmental studies of marine species. No formal web
but you can find some info at http://biology.caltech.edu/facilities/
Association of Marine Laboratories
(WAML): An association of the non-profit marine laboratories
by federal, state, university, and other non-profit organizations in
Western United States and Pacific Islands. Part of NAML,
the National Association of Marine Laboratories.
Institute of Oceanography:
Located in La Jolla, CA. Part of the University of
Diego. One of the west's eminent marine research laboratories.
Web Sites Focusing on Specific Marine Groups:
forum: A database
of information on nudibranchs, hosted by Bill Rudman of the Australian
Museum. Includes more than 30,000 images and facts on over
Recent and Fossil. Contains drawings, photos, and a
of bryozoan terms, links to bryozoan societies and sites, and on-line
regarding bryozoans. This site is maintained by Philip Bock
A web site
devoted to Anthozoans--taxonomy, experts in the field, species lists,
literature. Operated by Vreni Haussermann, from
specific information for the Pacific Northwest, but has taxonomic
Columbia: Has photos and descriptions of British
By Peter Egerton, a shell collector, web designer, and photographer.
of the world: Anemones, corals, cerianthids, and
A web site
based in the Netherlands and maintained by Arjan
photos and descriptions of a large number of ascidian species
Also includes links to other ascidian websites and to an ascidian
The goal of
this web site is to provide taxonomic information, keys, and
descriptions of the crustaceans of the world. Managed by an
coalition of taxonomists.
for United States: Home page for Ascidian news, a
by Charles and Gretchen Lambert on ascidians. Based in
and Fullerton, CA
This list is maintained by the C.B. Wilson Copepod Library at the
Museum, Washington, DC. It includes a bibliography of all
and branchiura literature, a world list of copepod and branchiura
and a list of specimens held by the Smithsonian Museum. The
list is now part of the World Register of Marine Species.
page: Claudia Mills is a specialist in gelatinous
based at University of Washington's Friday Harbor Marine
web site contains a lot of information on gelatinous species,
jellyfish. The site includes a list of errata
and Mills' 1998 book on Gelatinous Marine Zooplankton.
Database: Alpheids, especially genus Alpheus,
shrimp. We have no members of genus Alpheus
here in the Pacific
Northwest but we do have another genus in the family, Betaeus.
This database from the Smithsonian Tropical Research Institute and
by Arthur Anker has a general description of family Alpheidae and
many species in genus Alpheus.
Created and maintained by James Wood of the Bermuda Biological Station
for Research. Profiles a number of cephalopod species, has a
database on cephalopods, lists meetings on cephalopod biology, has
and movies of cephalopods.
Literature: This web site, part of the tree of life
is an assemblage of more than 7000 references on the systematics of
An increasing number, currently over 500, are available directly from
site as pdf files. The site is maintained by Dean Pentcheff,
Wetzer, and Joel Martin of the Natural History Museum of Los Angeles
CA. For the full site try this link: Decapoda.nhm.org
web site covers all living cephalopods. Created by James B.
and Catriona L. Day of Dalhousie University, it is housed at the
of Texas. CephBase provides taxonomic data, distribution,
videos, predator and prey data, size, references and scientific contact
information for all living species of cephalopods (octopus, squid,
and nautilus) in an easy to access, user-friendly manner.
Literature: A database of references on some of the
Arthropods in Infraorder Peracarida (specifically, the crustacean
Tanaidacea, Cumacea, and Mysidacea, and Lophogastrida).
by Gary Anderson at the University of Southern Mississippi
of the North American Pacific Coast: This web site
by Roger Clark
to the Thalassinideans
of the South Atlantic Bight (USA): This is a guide
to mud shrimp
(such as Upogebia)
and their relatives which live off the SE coast of the US.
document is available as a pdf at this link. Includes keys
Peer-reviewed and published by NOAA.
Zooplankton: A web site put together by Dave Wrobel
and Mills book).
by the Tromso museum, Norway.
Northwest Shell Club:
The web pages of this shell-collecting group contain photos of many
bivalve and gastropod shells and records of where the group has found
Amphipoda Database: Part of the MarineSpecies.org
Terms: Compiled by Joel Martin, Crustacean
curator, Los Angeles
of the Pacific Northwest: This site by Neil
McDaniel has great
photos and brief descriptions of about 30 seastars found off British
Many of the species are found in the Salish Sea, while others
are more northern species.
List of Marine,
Freshwater, and Terrestrial Isopod Crustaceans:
Brian Kensley, Marilyn Schotte, and Steve Schilling at the Smithsonian
Museum, Washington, DC.
site on gelatinous organisms (especially jellyfish) is maintained by
Wrobel. Dave has spent time as curator of jellies at Monterey
Bay Aquarium and New
Aquarium and is coauthor of the book Pacific
Coast Pelagic Invertebrates.
This web site focuses on taxonomy and descriptions of nemerteans,
coordinated by the International Congress of Nemertean
nemertean descriptions are very useful.
(Assembling the Tree of Life)
Decapoda: This site from the Los Angeles Museum of
contains thousands of article references, many available as pdf files,
on the systematics of decapod crustaceans. The site also
a list of decapod genera and a glossary of decapod biology.
web page. By Erik Thuesen, the author of the
(Light and Smith manual, Intertidal Invertebrates of Central
Hardy's Guide to Marine Gastropods: In 2018 contains a worldwide list
of about 60,000 gastropod and scaphopod species, with photos of about
23,000. Based on several collections, especially the Academy of
Natural Sciences in Philadelphia. Maintained by Eddie Hardy, who is
apparently a serious shell collector.|
Miscellaneous Web Sites:
Information System): From the USDA. Contains
on a large number of species found in the US, plus links to gene
PubMed articles, etc. A great source for information.
Columbia Marine Life: Photos of species from
various marine groups.
This web site is maintained by elasmodiver.
This site, produced by systematist Roderick Page at the University of
compiles a profile of what is known about species it is queried about
the fly". Information includes taxonomic information from
data, online images, and recent articles and abstracts from Google
A great resource!
Laboratory: Sidney, BC. A consulting
especially on marine sponges. For over 30 years Khoyatan
(KML) has been providing specialized consulting services on a wide
of marine topics. These include: Environmental Impact
including oil spills. Fishery Feasibility Studies, Species
Assessments of rare, threatened, and endangered species
2000: This extensive
database is also supported by ITIS. It is an attempt to
all species (animals, plants, fungi, and microbes). The
is based at the University of Reading, UK. Besides its own
species the site has links to many other "federated"
of May 2005, does not yet catalog invertebrates but it does include
Columbia Creature Page: Contains photos and brief
of many marine species in British Columbia. Many of the
can be found only by diving.
Dictionary of Invertebrate Zoology: A detailed,
and searchable dictionary in pdf format. Primary editor is
R. Maggenti from University of California, Davis.
This commercial web site of stock photos contains thousands of photos
marine species, along with many other types of photos. They
viewed for free but charge a fee of $9.00 or more for use.
affiliated site, Can
Stock Photos, offers many other photos, videos, and maps for
price or free.
Life: This web-based
project is an attempt to catalog the relationships among all living
Contains contributions from biologists from around the world, and
is composed of more than 3000 web pages.
This site, by diver and photographer Gary McCarthy, contains many
of underwater species, mainly from the warmer waters of southern
Hawaii, and the Caribbean.
Biological Indexer and Organizer:
(http://www.ubio.org) A digitized, online, searchable index of all
animal genera from Linnaeus down to the year 2004. Includes
of the complete version of the journal Nomenclator Zoologicus,
for the Zoological Society of London. This online version was
by David P. Remsen, Catherine Norton, and David J. Patterson,
Bull 2006 (210) 18-24
Publication Database of the Natural History Museum of Los Angeles
CA: A searchable archive of systematics references
and groups found along the Pacific coast.
for hundreds of thousands of digitized records of thousands of species
in museums across the US, including many marine species.
the National Science Foundation. Photographs of many of the
Another commercial web site that includes many good images of marine
such as sea slugs.
This educational web site by Thomas Carefoot of the University of
Columbia contains descriptions of the basic biology, photographs, and
of important scientific studies of a multitude of intertidal
found on the west coast of North America. The purpose is to
summaries of what is known about the invertebrates in order to
further undergraduate and graduate studies. The site's rich
currently includes thousands of photos and summaries of over 4000
papers. The organizing theme is a journey of a littorinid
subtidal water up to the upper intertidal.
Zoological Nomenclature: Specifies the rules for
such as these marine invertebrates.
The Ocean Biogeographic
Information System: A cooperative venture of the Alfred P.
Foundation, U.S. Office of Naval Resarch, National Science Foundation,
and National Oceans Partnership Program, Australia's CSIRO, Rutgers
and the Kansas Geological Survey. Provides biogeographic
from around the world. Can be searched by taxon or
Part of the 10-year Census
of Marine Life
site focuses on deep coldwater reefs from several regions of the world,
including the Aleutian Islands here in the NE Pacific.
photos, and descriptions of deep reefs.
Library: The British H.M.S. Challenger expedition
was one of the greatest oceanographic expeditions of all time and set
a surge of oceanographic studies around the world. It
80 volumes and over 30,000 pages of reports. A huge number of
marine species were discovered and described. This link to
of 19th Century Science, which is operated by volunteers, provides
to all the Challenger volumes. One may view individual pages
volumes on CD or DVD.
Library: Archives thousands of older or
biological books and articles and makes them available online (often as
pdf files). Examples include the entire 'Proceedings of the
States National Museum'. Also serves as the foundational
component of the Encyclopedia
database, based at the University of G?ttingen in Germany, specializes
in digitized versions of early species descriptions. It
links to more than 7000 digitized papers from 1500 on, including works
by Carolus Linnaeus. Nearly all the works relative to species
are complete up to 1770. Some references from later years are
included and the database continues to be updated. Searches can be made
by original taxonomic name, current taxonomic name, group, author, etc.
This web site,
based at the Smithsonian Environmental Research Center in Edgewater,
gives you access to a variety of databases which list invasive
species. Just enter your criteria and the
geographic area of
interest on the front page and the site will retrieve records from a
of relevant databases.
a world register of marine species. This web site is a
project by a worldwide group of marine taxonomists. It
databases of species names for a number of marine groups
sponges, several crustacean groups, pycnogonids, phoronids, and
Besides species lists it includes some depth and distribution
Portal: This European site is an initiative of ETI
under MARBEF (an EC funded network of Excellence) and KeyToNature (a
in the EC e-contentPlus Programme). In 2008 it incluced keys to 9875
and 5545 higher taxa, with descriptions and illustrations, synonyms and
vernacular names. It was contributed to by a wide network of
and is continuing to grow.
Library: This web site is an attempt by a
consortium of major
libraries such as the American Museum of Natural History, The Field
(Chicago), Harvard University botany libraries, Marine Biological
at Woods Hole Oceanographic Institution, Smithsonian Institution
etc. to make classical literature on biological species available
Documents include many classic books and journals, as well as a number
of recent works (example: Kensley
and Schotte, 1989. Guide to the Marine Isopod Crustaceans of
Caribbean, published by the Smithsonian Institution
photos and descriptions of over 225 species, covering all known
species except Epicaridea). Most are available as pdf files.
of Marine Species: This site is an attempt by a
number of universities
and government agencies to make a standard register of taxonomic names
and classification of marine species. Countries involved
many european countries, as well as South Africa and New Zealand as
as some participation by US scientists. The site is
taxonomists. Current "world lists" include groups such as
Pycnogonids, several groups of Crustaceans, and Ophiuroids.
has some more comprehensive regional lists as well, but these focus
on the Atlantic and Indian Oceans.
Contributions to Zoology: This entire line of
from the Smithsonian, many of which deal with marine species, is
Contributions to the Marine Sciences: This entire
in the 1970's) is available online, including a volume on Pacific
of the Southern California
Academy of Sciences: This journal frequently
about marine species along the Pacific coast of North
since 2000 can be found online.
of Life: A joint
project by a variety of scientists and organizations such as the
Woods Hole Biological Laboratory, Harvard University, the Field Museum
in Chicago, and the Biodiversity heritage library to make a database
a page for each known species on earth, with many links to different
of information, taxonomy, ecology, etc. Information is
many marine groups. This web site is a content partner to the
A web site
hosted by Florida State University which contains photos of thousands
species, including many marine species. Includes links to
The photos are available for "fair use".
Marine Life: This
international effort is sponsored by many organizations and government
agencies (spearheaded by the Alfred P. Sloan Foundation).
Washington, D.C., the 80+ nation organization has a 10-year initiative
to census marine species with emphasis on presence, distribution, and
for each species. The first comprehensive census is to be
This web site collects scientific names from a large number of other
including barcode of life, NCBI, GBIF, ITIS, etc. Its main
seem to be GBIF and EOL (Encyclopedia of Life). Any
add names to the index. Its intent is to become a list of all
names, whether correctly used or not. Inclusion in the index
not verify that the name is correct, but the index may provide links to
other resources that can help clarify the matter.
Information Facility: This site is an international,
collaboration for making biodiversity and bioinformatics information
available online. Its data portal allows one to search for
by species, by country, or by database.
site links to near real-time listening to a variety of deep-sea
distributed around the world. The site is run by the LIDO
(Listening to the Deep Ocean Environment).
Estuarine and Marine Exotic Specis Information System: This
created and maintained by the Marine Invasions Research Laboratory at
Smithsonian Environmental Research Center, provides comprehensive
on approximately 500 introduced marine and estuarine species of
and algae. The database can be searched by region or by
category. Currently the strongest emphasis is on species on
The Smithsonian Museum collection of nearly 19 million
Includes many photos, a bibliography, and a list of taxa for species
Antarctica to 30 degrees S.
on Orcas Island and sponsored by the UC Davis School of Veterinary
this web site focuses on many aspects of the ecosystem health of the
Juan Islands and Salish Sea. It includes information on
Sea species, accounts of ongoing research, maps of the Salish Sea
etc. A great site to visit for more information on the Salish
This site by the International
Commission on Zoological Nomenclature (ICZN)
is a repository for data on recent changes in zoological
is the official registry of zoological nomenclature and provides a
to register new nomenclatural acts, published works, and authors.
works in the region near the Rosario Beach Marine Lab to promote
education, and restoration projects and programs of the Northwest
Marine Conservation Initiative.
This web page, maintained by the Laboratory for Advanced Spatial
at the University of British Columbia, functions as an electronic atlas
for the wildlife of British Columbia, Canada, including marine
Each species listed has an accompanying atlas map showing where it has
been found in BC, plus references to other online databases where more
information can be found.
of Puget Sound:
This website is published by the University of Washington's Puget
Sound Institute. Its purpose is to be a central
source for integrated
scientific information about Puget Sound and the Salish Sea watershed
scientists, academics, and policy makers.
Explorer: This web site catalogs estimated
and level of endangerment for thousands of species in North
It makes estimates of status on many species that cannot be listed on
web sites because no formal status can be assigned to a species on a
web site unless some very specific studies (usually underfunded) are
International Barcode of Life:
A project with the goal of recording DNA sequences from all the living
species possible, in order to facilitate identification. A
database for iBOL is
and Scientists for Conservation (ASC): This
to outdoor extreme enthusiasts who want to go to exotic places to
environmental data for scientists. The organization pairs the
with the adventurers.
Seto Marine Biological Laboratory (P-SMBL):
Biological Laboratory is associated with Kyoto University,
It has been publishing in Marine Biology since 1949. Its
is the 'International Journal for Marine Biology'.
Sites Dealing with Introduced Species
along the Pacific Coast of America, especially the Salish Sea area:
The National Exotic Marine and Estuarine Species Information
This national database has a list of about 500 introduced marine and
invertebrate and algae species in the United States. Specific
reports are available only for tunicates, crabs, shrimp, and crayfish.
Council: Established by Executive order in 1999,
Federal departments and coordinates plans for dealing with invasive
Invasive Species page for
the Washington State Department of Fish and Wildlife:
plants and animals, freshwater and marine. Has a special
on invasive tunicates, along with descriptions of invasive tunicates in
Invasive Species: This web site covers more than
just New York.
It has incorporated the material from the former National Clearinghouse
for Aquatic Nuisance Species which was formerly maintained by Sea Grant
(but was closed because of budget cuts)
Project: Covers the entire Columbia River
freshwater and marine invasive species that may be found along the
of the Columbia River, including algae, Chinese mitten crabs, green
Atlantic Salmon, Spartina eelgrass, etc.
Maintained by Bonneville
Power Administration with input from US Fish and Wildlife, NOAA, and
Pacific States Marine Fisheries Commission.
Geological Survey NAS (Non-indiginous
Aquatic Species) site: Includes species lists for
Cnidarians, Crustaceans, Mollusks, and Vertebrates. No longer
updated because of budget cuts.
Center, Marine Invasions Research Lab: Based at the
Institution. Gary M. Ruiz, Senior Scientist.
Aquatic Invasive Species
Network (CAISN), a part of NSERC (Natural Sciences and
Research Council), Canada.
Introductions into British Columbia Marine Waters, Major Trends:
Part of E-Fauna BC, The electronic atlas of fauna of British
Contains a list of known marine introductions in British Columbia.
Invasive Species Page by Fisheries and Oceans
photos and links to information and booklets on a number of marine
species found in Canada.
Invasive Species Lab:
Lab of Evgeny Pakhomov, University of British Columbia, Vancouver,
Contains information on projects the lab is doing and on BC invasive
Hotline: To report suspected invasive species
discovered in Oregon.
Has databases on plants and animals, mostly terrestrial and freshwater.
a site maintained by CABI.org, a not-for-profit international
Some of the marine species pages seem not well developed yet, but can
some information on species invasive internationally.
Invasive Species Database: 100 of the World's Most Invasive
Species. Includes terrestrial, aquatic, and marine
Species Page of the Western Ecology division of the United
Protection Agency (EPA): Links to a large pdf atlas listing
and estuarine ononindiginous species in the western US, along with
and habitat information for each species. Also links to a
Access file, PICES, which was used to generate the atlas.
Exotics Guide: Non-Native
Marine Species of the North American Pacific Coast:
site, by Andrew Cohen and produced by the Center for Research on
Bioinvasions of the San Francisco Estuary Institute, contains photos
descriptions of many exotic species. San Francisco Bay is one
the main centers for aquatic invasive species on the west coast, so
site should have a firsthand look at many of them. The site
attempts to catalog species found elsewhere along the Pacific coast.
Red List: This
is a list of the endangered status of species around the world by the
Union for the Conservation of Nature (IUCN). For each species
IUCN assessment is given, along with an explanation of why it is
(including because of competition from introduced species)
Crab web pages from University of Washington College of the
for Washington State Sea Grant: Green Crabs threaten to
in Washington State waters. This web site details for
to identify them and whom to contact if they are found.
Return to Main Inverts Key
Salish Sea Invertebrates web site provided courtesy of Walla